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Glucose or fructose (10 mM), oligomycin (2 µM), and 2-DG (50 mM, all from Sigma-Aldrich) were sequentially added into the cells during real-time measurements of extracellular acidification rate (ECAR) and OCR (Oxygen consumption rate) using an XFe24 analyzer (Seahorse Bioscience)

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It is important to note that sulfate is the rate-limiting factor in both PAPS and GAG synthesis, meaning that sulfate insufficiency will reduce the synthesis of both products